This
tissue wraps-in and separates muscle fibres, acts as supporting material,
and helps movements, protects muscles against shocks and too important tension.
Scientific
name of the muscle fibre. Mutiplied thousands of times, it makes up
the whole muscle. It is wrapped around by a protective girdle
called conjunctive tissue.
They are made of fibrils, constituted of thin threads themselves made of
proteic molecules.
Their size (at adult age) ranges from 10 to 100 angstrom in diameter ( with
up to 20cm length). A the origin, during the very first stages of embryo
development, muscle fibres develop out of undifferenciated cells calles myoblasts,
which merge into one another. After birth these grow in size but not in number.
There is a possibility to regenerate them after an injury, out of other undifferenciated
cells called "satellite cells". Yet, generally speaking, if the
loss of muscle fibre is too important, it is thanks to hypertrophia (increase
in size)
that compensation occurs.
This
refers to the sub-partition of cells. It encloses and separates a volume
from the cytoplasm. In certain types of muscle fibres, the endoplasmic reticulum
of the "smooth" type used to store calcium is
called "sarcoplasmic reticulum" (found in muscle cells). It is
in charge
of storing and releasing calcium.
They
are parallel to one another, and laid in the same direction. A Myofibril is
made of a succession of sarcomeres which
are the minimal muscular contraction unit.
These sarcomeres are
laid in chains on the whole length of the fibre.
This
refers to the volume situated between Myofibrils and
the endoplasmic reticulum in which calcium is
released by the sarcoplasmic
reticulum wrapped around Myofibrils.
Cytosol only contains
soluble substances. Water amounts generally to around 85
% of cytosol contents. With its
suspended macromolecules in a aqueux and salty environment, it has
a viscosity four times more important than that of water and this corresponds
to that of a celloïd gel.
Sarcoplasmic
reticulum is the name given to the smooth endoplasmic reticulum found in
striated skeletal muscle cells. This zone is a store of Ca2+ ions (hence
is it called
a "calciosome") which are released into the cytoplasm in reaction
to a plasma membrane depolarisation,
caused by the binding of ACh on muscle cells nicotinic receptors. This increase
in cytoplasmic calcium enables
the interaction between myosin and actin threads. Calcium is
then pumped (thanks to a "pump" using ADP to operate) back to the
sarcoplasmic reticulum.
These
are organites in charge of breathing: filled with enzimes which transform
sugar components and fat into ATP thanks
to oxygene. They thus oxydate carbon and glucose to provide energy. It is
at that level that carbon oxydation reactions
occur which produce ATP.
It is a that level that occurs glocose oxydation too.
Glucose turns into tiny carbonate
molecules called « pyruvate ». Pyruvate is then carried into
the mitochondrion matrix, where it is gradually deprived of its carbon and
hydrogene components to then create, among many other things, ATP and
CO2.
This
is the minimal unit involved in muscular contraction. There is not length
modification involed during a contraction but a gliding of threads one along
the other, which creates a general contraction. The sarcomere's
length is reduced by a roughly 20% margin. The speed of contraction is 15
angstrom/second.
The more a muscle is divided into sarcomeres, the
more it will give the impression of shortening.
During the shortening phase of each sarcomere there
is no shortening of either thin or thick threads, but a gliding motion leading
them to overlap one another. This could be compared to a rowboat. At the
core of each sarcomere is found the myosin molecules,
each surrounded on both sides by six actin molecules.
The myosin molecule
is going to connect to actin outstretching
its "arms" and draw them inwards. At the very end of the sarcomère (on
both sides)
is an area called "Z Line" which enables actin molecules
to connect, thus ensuring a connexion between each sarcomere with
both the following and preceeding one.
The
tip of myosin molecules
moves forward on a distance approximatively corresponding to the diametre
of the actin molecules
thread. This bending is made possible by the hydrolysis of
an ATP molecule which binds onto the
tip of myosin.
Each myosin molecule
is endowed with a "tip" or "tail" running along its axis and fitted with
two globular "heads" (crossbridges).
Two globular heads are thus ready for bindings: one for the binding of ATP and
the other for the binding onto actin.
The crossbridge site is
located on each side of the tip of each myosin molecule
and draws actin molecules
threads inward and rearward, thus shortening the whole sarcomere.
The actin molecule
is a globular molecule which combines to others to make up a chain. This
chain has a coiling pattern, and constitutes what is known as a "thin" thread.
Each actin molecule
owns a binding site for myosin.
A thin thread owns thus many binding sites. An actin molecules
thread homes two other ones: the tropomyosin molecule
which coils around it (and, by doing so, physically inhibits the actin's binding
sites) and troponin molecules
(of globular shape) acing as connexion between actin and tropomyosin For
muscle contraction to begin, the troponin molecule
coiled around actin must
move and thus drag the tropomyosin molecule
away from the binding sites on which myosin will
bind. For troponin molecules to move calcium has
to fix itself onto them. When this is done, the overall shape of troponin molecules
is modified and this one alters its position, hence dragging tropomyosin along,
which in turns uninhibits the actin binding
site, in turn enabling myosin to
get in contact with it. When calcium leaves troponin molecules,
they get back to their original location and "push" tropomyosin back
in front of binding sites, and thus prevents muscular contraction to take place.
Globular
in shape, it is both fixed to tropomyosin (coiled
around actin, and
right on its binding sites) and to actin. Troponin molecules
function as ball-bearings bearing on a larger "cable" (tropomyosin"
to make it move on actin.
Tropomyosin is
usually coiled over most of the actin's
binding sites, thus making impossible any contact from myosin's crossbridging
sites
to actin, and thus
preventing muscular contraction to occur.
Troponin is thus the molecule which,
thanks to its movements, indirectly uninhibits or inhibits the actin 's
binding sites by altering the position of the coil of tropomyosin.
We are here facing a proces similar to that of a latch. When a set of calcium ions
fix themselves onto troponin the
latter rotates (such as the cylinder of a lock) and this movement drags tropomyosin away
(such as a latch that once kept the door closed), and then uninhibits the
binding site for myosin's
molecules' crossbridging.
This
is a long coiled molecule acting as a muscular contraction lock. This molecule
is coiled around the thin thread molecule, and is moved away by the motions
of another molecule to which it is bound. Tropomyosin can
also be considered as the biological equivalent of a plug-mask or a latch,
or more simply, a bit of plastic put on a magnet to prevent connection.
This
is the name given to the end part of myosin molecules,
involved in a binding-deconnecting cycle with actin binding
sites, leading to the shortening of sarcomeres and
thus to the creation of power. This crossbridging
site needs
ATP to function, and uses it to find
the energy required to move and fetch the actin 's
binding site, then uses another ATP molecule
again to unhook from it. This is what is known as the crossbridging
cycle : the process is as follows: binding of myosin's
crossbridging site onto actin's
binding sites,
rearward movement, unhooking of the crossbridging
site, followed by a forward movement from it to fetch another actin's
binding site, etc.
This
is the area where connection between myosin's
molecules' crossbridging sites and actin occurs.
It is at that level that myosin binds
to actin to
then draw it rearward, thus shortening the sarcomere and
creating a pull-force. These actin's
binding sites are generally inhibited by a molecule called tropomyosin,
coiled around actin,
which makes muscular contraction impossible as long as it has not withdrawn.
This
refers to the division of myofibre corresponding
to the sarcomere.
This
refers to the central zone of sarcomeres :This is
the area where one can only find myosin,
the area towards which actin threads
will be drawn during muscular contraction. Each myosin molecule
is surrounded by actin molecules,
and when these are drawn inwards by the action of myosin this
movement occurs in the direction of myosin's
center, or "H-line".
This
is the opposite zone to that of the "H-line", that is to say the area where
only actin molecules
are to be found, at the end of sarcomeres; it is at that level that inward
movements begin.
This
is the area between two sarcomeres :
it is at that level that muscle stretching occurs. When a muscle stretches
because of a weight beyond its contractile power, sarcomeres do
not stretch, but the only "elastic" zone does, and it is that between
them, called "Z-line".
Calcium is
a fundamental ion for cell functionning. It is indirectly responsible for
the "opening" of
the actin molecules
binding site, enabling myosin molecules
to bind to them. This opening happens when calcium fixes
itself onto troponin and, by
doing so, alters its general shape, in turn forcing troponin to
settle back, which in turn again trigers a change in tropomyosin molecules
position, that previously inhibited that actin's
binding site. Calcium is released
from stores located around Myofibrils (sarcoplasmic
reticulum). It gets fixed
on troponin thanks
to electrical phenomena.
Lactic
acid is the by-product of muscle glycogene consumption. Glycolysis (sugar
muscle consumption to create energy) can either be done in a sufficient/non-sufficient
oxygene environment (aerobic/anaerobic environment). In the case of aerobic
glycolysis this consumption of sugar occurs at the level of mythochondrions,
which will
completely consume lactic
acid indirectly produced. In the case of an effort without enough oxygene
provided, sugar consumption occurs into the cytoplasm and lactic acid can
not be eliminated thanks to perspiration.
When
an ATP molecule is hydrolysed (dissolved
into water) it "breaks apart" into an ADP element on the one hand and into
a Phophate element on the other hand, and releases energy. The coupling back
of these two elements makes the regeneration of ATP possible,
but it will require the same amount of energy to do so.
Molecule
with very energetic atomic links. This is a very energetic phosphate compound.
This molecule is used to produce energy. Its breaking up by water (hydrolysis)
releases energy (around 30kj/mol), and its synthesis consumes the same amount
of energy. ATP Hydrolysis gives an ADP molecule and a phosphate compound.
Action
of water cutting molecules and releasing chemical energy (in the case of ATP hydrolysis).
The basic idea is to disrupt molecules to release energy.
occurs
when a compound such as ADP becomes ATP.
This is the reverse phenomenum of hydrolysis. Energy is here used to merge
instead of disrupting.
When ATP loses
an electron, captured by another molecule.
Either
loss of electrons (becomes positive) or electron addition (becomes negative).
Transmitting
the message of Calcium release.
These are nerve cells the axons of
which divide into branches connecting to a maximum of myofibrils.
The more axons within myofibres receiving calcium release
messages, the more sarcomeres contractions will
occur, and so the most powerful will contraction be.
This
is the unprotected end of motor neuron (unprotected by any girdle), the one
in charge of transmitting information via a neurotransmeter. As an electric
cable, it corresponds to the end part, where the wires are left unprotected
for connection.
The
Motorneuron + myofibre unit is
refered to as a « motor
end plate ». This
is the zone on which connection between the nervous system and the muscular
system occurs. At the level of myofibres,
in the groves of the plasma membrane surface (protecting Myofibrilles)
we can find a « motor
end plate » where motor axons bind. At the level of axons ends
are vesicles releasing a neurotransmetter (Ach).
When the action potential binds to the motor
end plate it depolarises the
plasma membrane thus opening the ducts releasing calcium.
These ducts are said to be "potential-dependant", because they
need an action potential to move. This calcium hence
released will enable the release of a neurotransmetter called Ach binding
onto the motor end plate and
transmitting an action potential again. The motor
end plate carries an enzyme (choline) which will use the connection to
go up the axon end and
be used to create another Ach. As
a home electric connection, this would correspond to the electric "contact" parts.
This
is the name given to the MotorNeuron--myofibre system:
since each motor-neuron is
connected to a muscle fibre, the complete muscular contraction is the fruit
of the simultaneous action of these independant networks of connection: many motor-nerons will
simultaneously stimulate the myofibre to
which they are linked and the sum-total of these stimulation-contraction
phases of each single motor-unit will create power. Compated to an internal
combustion engine, the motor
unit would roughly correspond to the combustion chambre - sparking plug
unit; the sparking plug is the motor-neuron bringing up information, and
the myofibre would be the combustion
chamber where chemical reactions occurs, leading to the creation of power.
The simultaneous involvement of each plug-chamber system (motor
unit)
creates our movements.
This
is what is usually refered to as "nerve influx", which is in fact
made up of a series of action potentials. This corresponds to a change in
cell polarity.
These are ions (potassium,
sodium) which are at the origin of such electric potential difference between
the inside and the outside of a cell. The action
potential is made up of a series of events: first occurs a short-lived
and local depolarisation,
followed by an increase in internal membrane potential, and finally by a
repolarisation of the internal membrane. An action
potential lasts between 2 and 3 milliseconds.